Microbial
Ecology - Biology 4446/5446 (3-4)
Spring
2004
Instructor: Dr. R.J.C. McLean, Dept. Biology,
Office: SCI 364, office hours M-Th 8:
Prerequisite: Bio 2400/3440, courses in biochemistry and/or
ecology would be helpful
Text: R.M. Maier, I.L. Pepper, and C.P. Gerba 2000. Environmental
Microbiology, Academic Press,
Other reference materials: (Old text) R.M. Atlas and R. Bartha. Microbial
Ecology, Fundamentals and Applications, 4th. ed., 1997.
Benjamin/Cummings Publishing Co. Inc., Menlo Park CA. Scientific journals
including: Applied and Environmental Microbiology, Advances in Microbial
Ecology, Environmental Microbiology, FEMS Microbiology Ecology, Microbial
Ecology, Microbiology and Molecular Biology Reviews, Annual Review of
Microbiology
Catalog Description: This course will illustrate the wide variety of
bacteria in nature, their interactions with other organisms and environments,
and their roles in global cycling of elements such as carbon, nitrogen, and
sulfur. The laboratories involve a semester-long project dealing with
Course grading:
Mid
term 1 (Feb 24) 10 %
Mid
term 2 (April 8) 15 %
Laboratory 50 %
Final
take home exam 25 % (Due at
The laboratory grade (50 % of course grade) is broken down as follows:
Laboratory quiz based on class experiments 10 %
(last week of classes)
Major laboratory project (aquifer/terrestrial microbiology) 40 %
(research proposal 5%, oral
presentation 10%, effort 5%, report 20%)
5446 students will be required to submit a budget containing detailed
costs of all supplies at the time of their proposal.
As a guide, I use
the following grading scheme: [A 90-100%, B 80-89%, C 70-79%, D 60-69%, F <
60%]. In situations where a student's
grade is within 2% of the cutoff (e.g. 88-92), the final grade will be based on
both the grade achieved, and the lab grade for effort and participation. Grade
disputes must be given to Dr. McLean in writing within one week after a test is
returned. This will result in the entire test being regraded and the higher or
lower grade being used.
** Students must achieve a passing grade (at least 60%) in both
the lecture (mid-term 1, mid-term 2 and final) and laboratory portions of the
course in order to pass. Students dropping the course before March 8 will get
an automatic W. After March 8, students must have a passing grade in order to
drop with a grade of W, otherwise a grade of F will be assigned.
Subjects Covered:
A. Overview of Environmental Microbiology
Methods employed
Microbial Communities
Physical and Chemical
Influences
Biological interactions
Biogeochemical cycles
B. Water Microbiology and Aquatic Environments
C. Terrestrial and Subsurface Microbiology
D. Extreme Environments and Life on other worlds (exobiology)
Laboratory Topics:
(Many of these exercises will run over several weeks)
1) Direct examination of microbial populations in the
environment (bright field, dark field, fluorescence microscopy)
2) In situ determination of microbial metabolism in the
environment (CH4 production by sediment microorganisms)
3) Enrichment culturing for selected groups of microorganisms (e.g.
autotrophs, heterotrophs, aerobes, and anaerobes)
4) Microcosm and mesocosm studies i.e. modelling an environment
5)
Microbial Evolution (spread of
antibiotic resistance in a microbial population)
6)
Coliform testing
7)
Major Laboratory Project.
The major laboratory
project will consist of students investigating some aspect of microbial
ecology. In many cases, students may elect to examine the Edwards Aquifer, or
alternatively another project with approval of Dr. McLean. The fascinating
aspect of most of these studies is that they will represent original research.
Students will work on these projects in pairs. Some possible projects include:
1)
Phenotypic characterization of
mutants in Myxococcus
2)
Characterization of an
unidentified new aquifer isolate.
3)
Estimating decomposition rates
of various materials (e.g. leaves, insects, gasoline) by aquifer microorganisms.
4)
Influence of artificially
introduced wastewater on the aquifer microflora (fate of introduced E. coli or other organism).
5)
Do aquifer bacteria assist in
mineral formation or dissolution.
6)
Influence of bacterial biofilms
on leaf colonization by macroinvertebrates such as snails.
7)
Chemotaxis response of aquifer
isolates to acyl homoserine lactones (AHLs).
8)
Enhancing biofilm formation in E. coli UTI strains for genomic
investigation.
9)
AHL antagonists in plants or
invertebrates.
10)
Influence of organic or
inorganic pollutant on aquifer bacteria - can also involve experimental
estimation of bioremediation potential of aquifer bacteria.
11)
Isolation of unusual
microorganisms (e.g. anaerobes, methylotrophs).
12)
Your own idea.
Students working in
pairs will select their topic and get approval from Dr. McLean by the end of
the first week of classes. During the second week of classes (lecture and lab),
students will turn in 2 page typed research proposals and present (5 min oral
presentations) research proposals consisting of: problem, background, proposed
strategy to be used for experimentation, data analysis techniques, and
anticipated findings. Participation in discussion by all members of class is
required.
The final report for
this project will be presented both as an oral report (10 minute presentation
and 5 minute question) and as a written scientific paper (using the format for
Applied and Environmental Microbiology available at http://aem.asm.org/misc/ifora.shtml).
The paper is due the last day of classes (